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Houthis, who have held Sanaa and much of northern Yemen for years, seized Mokha and key Red Sea islands this month.

Saudi-led coalition said it intercepted a dawn Houthi attack, while some Riyadh residents reported hearing an explosion.

Deadly clashes erupted in Yemen on Thursday, after Houthi forces began a ground assault accompanied by rocket and drone attacks.

A Houthi leader said Saudi-backed troops were targeted in the attacks and that "hundreds" were either killed or injured.

The rebels also raided U.N. offices in Sanaa on Aug. 31 and detained 19 employees, according to the U.N.

Major General Mohammed al-Ghamari died alongside "companions" and his 13-year-old son, the Huthi statement said, without giving the date of the attack.