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A Houthi leader said Saudi-backed troops were targeted in the attacks and that "hundreds" were either killed or injured.

The rebels also raided U.N. offices in Sanaa on Aug. 31 and detained 19 employees, according to the U.N.

Major General Mohammed al-Ghamari died alongside "companions" and his 13-year-old son, the Huthi statement said, without giving the date of the attack.

The Houthis separately threatened to “employ all means and instruments available” to target several American oil firms.

Medics were still searching for victims believed to be trapped under rubble, the rebels said, indicating the casualty tolls could rise.

The attack happened about 225 kilometers (140 miles) off the coast of Aden, which is held by forces loyal to Yemen’s exiled government.