Mahant Dinendra Das said the "entire fault" lay with Gopal Rao Nagarkatte, whose role in overseeing temple operations and donation management is also under scrutiny.
Addressing a press conference alongside SP chief Akhilesh Yadav, Hari Shankar Safariwala, the head of the Ram Niwas temple, alleged that his complaints against Rai were ignored by the authorities.
Investigators believe the recovered "Ramrajya Kosh" chest could prove to be key evidence in establishing the alleged modus operandi and identifying everyone involved.